CLASSIFICATION OF VERONICEAE

This classification of Veroniceae is mostly accomplished by Dirk Albach and Montserrat Martínez-Ortega and still preliminary. It shall here give a provisional overview of our understanding at the moment, which taxa should be recognized.

The following genera are included in Veroniceae. Click on the name to see its position in our classification:

Beccabunga,Besseya,Botryopleuron,Calorhabdos,Chionohebe,Cochlidiosperma,Derwentia,Detzneria,Falconeria, Gymnandra, Hebe, Heliohebe, Kashmiria, Kokoneria, Lagotis, Leonohebe, Leptandra, Neopicrorhiza, Oligospermum, Paederota, Paederotella, Parahebe, Picrorhiza, Pseudolysimachion, Scrofella, Synthyris, Veronica, Veronicastrum, Veronicella, Wulfenia, Wulfeniopsis
 
 

Scrofella Maxim. Bull. Acad. Imp. Sci. Saint-Petersbourg 32: 511. 1888.
 
 

Type: Scrofella chinensis Maximowicz (monotypic genus)
 
 


 
 

Figure from: Flora of China, Illustrations, Vol. 18, Zhengyi, W. & P. Raven. 2000. Missouri Botanical Garden Press.
 
 

Veronicastrum Heist. ex Fabr. Enum. Meth. Pl. Hort. Helmstad. 111. 1759. – Type: Veronicastrum virginicum (L.) Farwell

Comprises:

Botryopleuron Hemsl. Hook. Icon. Pl. 27: t. 2670. 1900.

Calorhabdos Benth. Scroph. Ind. 44. 1835.

Veronicastrum sect. Calorhabdos (Benth.) Hong Fl. Reip. Pop. Sin. 67(2). 401. 1979.

Leptandra Nuttall. Gen. N. Amer. Pl. 1: 7. 1818.

Preliminary list of species:

Veronicastrum axillare, brunonianum, caulopterum, cerasifolium, formosanum, kitamurae, latifolium, liukiuense, longispicatum, lungtsuanense, plukenetii, rhombifolium, robustum, sibiricum, simadai, stenostachyum, tagawae, tubiflorum, venosum, villosulum, virginicum, yamatsutae, yunnanense
 


Photo: Veronicastrum virginicum, D. Albach, BG Wien

Lagotis Gaertn. Novi Comment. Acad. Sci. Imp. Petrop 14: 531-537. –

Type: Lagotis glauca Gaertn.

= Gymnandra Pall. Reise durch verschiedene Provinzen des Russischen Reichs, Vol. 3. Kaiserliche Akademie der Wissenschaften, St. Petersburg, p. 710.

= Kokoneria Keng & Keng f. J. Wash. Acad. Sci. 35(12): 374-378.
Comprises:

Lagotis sect. Caulescentes Maxim. Bull. Acad. St. Petersb. 27: 523. 1881

Lagotis sect. Acaulescentes Maxim. Bull. Acad. St. Petersb.

Lagotis sect. Schizocalyx Tsoong. Fl. Reip. Pop. Sin. 67(2): 327. 1979
 
 

Preliminary list of species:

Lagotis alutacea, angustibracteata, brachystachya, brevituba, cashmeriana, clarkei, crassifolia, decumbens, glauca, globosa, hultenii, humilis, ikonnikovii, integra, integrifolia, kongboensis, korolkowii, kunawurensis, macrosiphon, minor, pharica, praecox, ramalana, reniformis, stelleri, stolonifera, uralensis, wardii, yunnanensis
 
 

Kashmiria Hong Bot. Not. 133: 565-567. – Type: Kashmiria himalaica (Hook. f.) Hong (monotypic genus)

= Falconeria Hook f. Ic. plantarum 15: t. 1483. 1883.

= Wulfenia auct. non Jacq. Pennell. Acad. Nat. Sci. Philadelphia Monogr. 5: 67. 1943.

Wulfenia Jacq. Misc. Austr. 2: 60. 1781. – Type: Wulfenia carinthiaca Jacq.

Preliminary list of species:

Wulfenia carinthiaca, W. baldacii, W. orientalis
 


 

Photo: Wulfenia orientalis, D. Albach, BG Bonn
 

Picrorhiza Royle ex Benth. Scroph. Ind. 47. 1835 – Type: Picrorhiza kurrooa Royle.

Comprises:

Neopicrorhiza Hong. Opera Bot. 75: 56. 1984.

Preliminary list of species:

Picrorhiza kurrooa, P. scrophulariiflora, P. minima
 


 

Wulfeniopsis Hong Acta Phytotax. Sin. 18: 50-52. 1980.– Type: Wulfeniopsis amherstiana (Benth.) Hong

=Wulfenia auct. non Jacq. Benth. Scroph. Ind. 46. 1835.

Preliminary list of species:

Wulfeniopsis amherstiana, W. nepalensis
 
 

Paederota L. Spec. Pl. ed. 2: 20. 1762. – Type: Paederota bonarota (L.) L.

Preliminary list of species:

Paederota lutea, P. bonarota

Photo: P. bonarota, Dirk Albach, South Tirol
 
 

Veronica subgen. Hebe (Juss.) Martínez-Ortega, Albach & Fischer (ined.) – Type: Hebe magellanica Gmelin Syst. Nat. 2: 27. 1791.

= Hebe Comm. ex Juss., Gen. Pl.: 105. 1789 [basionym]

= V. sect. Hebe (Juss.) G. Gon, Gen. Hist. 4 (2): 570. 1838.

Comprises:

Chionohebe B. G. Briggs & Ehrend. in Contr. Herb. Australiense 25: 1-4. 1976. page?

Detzneria Schltr. ex Diels. in Bot. Jahrb. Syst. 62: 490. 1929.

Hebe Comm. ex Juss., Gen. Pl.: 105. 1789

HeliohebeGarnock Jones, New Zeal. Journ. Bot. 31: 323. 1993.

ParahebeW. R. B. Oliver in Rec. Domin. Mus. 1: 229. 1944, excl. P. litophylla B. G. Briggs & Ehrend.

 Leonohebe Heads. Otago Bot. Sco. Newslett. 5: 4-11. 1987.

Preliminary species list will be assembled by Dr. Phil Garnock-Jones, New Zealand.

Photo: V. macrantha, Dirk Albach, RBG Kew

 

Veronica subgen. Derwentia (Raf.) Martínez-Ortega, Albach & Fischer (ined.) –
        Type: V. derwentia Andrews

= V. [unranked] Calycinae Benth. in DC., Prodr.: 477. 1846. [basionym]

= V. [unranked] Calycinae (Benth.) Römpp in Repert. Spec. Nov. Regni Veg. Beih. 50: 135. 1928.

= V. sect. Veronica pro parte

= DerwentiaRaf. Fl. telluriana pt. 4. 1836.

Comprises among other species:

V. sect. Labiatoides Wettst. in Engler & Prantl, Nat. Pflanzenfam. IV, 3b: 86. 1891

V. sect. Veronica subsect. Calycinae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977

Parahebe litophylla B. G. Briggs & Ehrend. in Telopea 5 (1): 255. 1992.
 
 

 Preliminary list of species:

Veronica arcuata, arenaria, arguta, blakelyi, brownii, calycina, decorosa, derwentiana ssp. anisodonta, ssp. homalodonta, ssp. maideniana, ssp. subglauca, ssp.derwentiana , distans, formosa, gracilis, grosseserrata, hillebrandii, monantha, nivea, notabilis, Parahebe lithophila, parnkalliana, perfoliata, plebeja, sobolifera, subtilis, velutina
 

Photo: Veronica derwentiana, from I. R. McCann. 1992. The coast and hinterland in flower. Victorian National Parks Association.

 

Veronica subgen. Pseudolysimachion (W. D. J. Koch) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. spicata L. Alternative: Hedystachys Dumort.

= V. sect. Pseudolysimachium [Pseudo-Lysimachium] revise art. 60 ICBN W. D. J. Koch, Syn Fl. Germ. Helv.: 527. 1837 [basionym] =Pseudolysimachion (W. D. J. Koch) Opiz, Seznam: 80 (1852) Comprises:

V. sect Pseudolysimachium subsect. Longifoliae (T. Yamaz) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 151. 1977.

V. sect Pseudolysimachium subsect. Alatavicae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 151. 1977.

V. sect Pseudolysimachium subsect. Spuriae (Holub) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

V. sect Pseudolysimachium subsect. Dahuricae (Holub) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

V. sect Pseudolysimachium subsect. Pseudo-Lysimachium (W. D. J. Koch) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

V. sect Pseudolysimachium subsect. Pinnatae (Holub) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

V. sect Pseudolysimachium subsect. Schmidtianae (T. Yamaz.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

Photo: Veronica longifolia, Dirk Albach, Bot. Gard. Genève

Preliminary list of species:

 Veronica alatavica, arenosa, bachofenii, barrelierii spp. andrasovszkyi and spp. barrelierii, crassifolia, dahurica, euxina, incanaspp. incana and spp. pallens, insulare, kiusiana, komarovii, linariifolia, longifolia, maritima, miyabei, nakaiana, ogurae (ined.), olgensis, orchidea, ornata, ovata, pinnataspp.laeta and spp.pinnata, pyrethrina, rotunda spp.subintegra and spp.rotunda, sachalinensis, sajanensis, schmidtiana, senanensis, sieboldiana, spicataspp. fischeri, spp. lanisepalum, spp. porphyriana, spp. spicata and spp. viscosula, spuria, subsessilis
 

Veronica subgen. Beccabunga (Hill) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. beccabunga L.

= Beccabunga Hill, Brit. Herb.: 94. 1756 [basionym]

= V. sect. Beccabunga (Hill) G. Don, Gen. Hist. 4 (2): 576. 1838.

= V. sect. Veronicastrum W. D. J. Koch, pro parte

= V. sect. Pocilla Dumort., pro parte
 
 

 Comprises: V. sect. Beccabunga subsect. Beccabunga Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Alsinebe subsect. Peregrinae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.

V. sect. Veronicastrum subsect. Veronicastrum (W. D. J. Koch) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977.

V. sect. Veronicastrum subsect. Serpyllifoliae (G. Don) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977. (incl. V. archboldii Pennell in Brittonia 2: 182. 1936.)

V. sect. Veronicastrum subsect. Telephiifolia Mart. Ortega & E. Rico in Bot. Journ. Linn. Soc 135: 186. 2001. [=V. sect. Veronicastrum subsect. Minutae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977, pro parte, excl. V. minuta, V. chionantha, V. kopetdaghensis et V. violifolia].

V. sect. Alsinebe subsect. Acinifolia (E. B. J. Lehm.) Stroh in Beih. Bot. Centralbl. 61 B: 398. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.
 
 

 Preliminary list of species:

Veronica acinifolia, americana, anagallis-aquatica ssp.anagallis-aquatica, ssp.lysimachioides, ssp.oxycarpa, anagalloides, archboldii, balansae, beccabunga ssp. abscondita, ssp.beccabunga, ssp. muscosa, bozakmanii, catenata, charadzeae, colchica, davisii, debilis, gentianoides, glaberrima, gorumsensis, heureka, hispidula, kaiseri, kemulariae, kopgecidiensis, langei, michauxii, nudicaulis, oetaea, peregrina ssp.peregrina, ssp.xalapensis, perpusilla, platycarpa, poljensis, pusilla, quezelii, repens, reuteriana, riederiana, scardica, schistosa, serpyllifolia, sumilensis, syriaca ssp.aksekiensis, ssp. syriaca, telephiifolia ssp.glareosa, ssp.telephiifolia,tibetica, undulata,

Photo: V. beccabunga, Dirk Albach, Swiss Alps

 

Veronica subgen. Veronica L., Sp. Pl. 9. 1753. – Type (see Britton & Brown, Ill. Fl. N. U. S. 3: 199. 1913): V. officinalis L. – Lectotype (designated by Sutton in Jarvis & al., A list of Linnean generic names and their types. International Association for Plant Taxonomy: 98. 1993): Herb. Clifford 8, Veronica 4 (BM).

= V. sect Veronica pro parte

= V. sect. Veronicastrum W. D. J. Koch, pro parte

Comprises:

V. sect. Beccabunga subsect. Scutellatae (G. Don) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Montanae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Veronica Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Javanicae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977, pro parte [excl. V. javanica].

V. sect. Veronica subsect. Canae (T. Yamaz.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Pyroliformis Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Szechuanicae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 153. 1977.

V. sect. Veronica subsect. Urticifoliae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977.

V. sect. Veronicastrum subsect. Gouani (Moretti) Stroh in Beih. Bot. Centralbl. 61 B: 385. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977, pro parte [excl. V. monticola, V. lanosa, V. mexicana].

V. sect. Veronica subsect. Carpathicae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Veronica subsect. Aphyllae (Römpp) Stroh in Beih. Bot. Centralbl. 61 B: 425. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Veronicastrum subsect. Glandulosae (Römpp) Stroh in Beih. Bot. Centralbl. 61 B: 388. 1942.

V. sect. Veronicastrum subsect. Alpinae (Benth.) Stroh in Beih. Bot. Centralbl. 61 B: 386. 1942., pro parte [excl. V. chinoalpina]

Photo: V. alpina, D. Albach, Vorarlberg, Austria
 

Preliminary list of species: Veronica abyssinica, allionii, alpina, aphylla, baumgartenii, bellidioides, cana, chayuensis, copelandii,cusickii, dabneyi, deltigera, fargesii, glandulosa, grandiflora, gunae, henryi, japonensis, laxissima, longipetiolata, mannii, miqueliana, montana, morrisonicola, muratae, nipponica, officinalis, oligosperma, onoei, piroliformis, ponae, riae, robusta, scutellata, stelleri, sutchuensis, szechuanica, taiwanica, tsinglingensis, uralensis, urticifolia, vandellioides, wormskjoldii, yunnanesis
 
 

Veronica subgen. Orientales (E. Wulff) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. orientalis Mill.

= V. [unranked] Orientales E. Wulff in Trudy Tiflissk. Bot. Sada 15: 59. 1915. [basionym]

= V. sect. Veronica pro parte

Comprises:

V. sect. Veronica subsect. Pentasepalae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Caucasicae (Riek.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977.

V. sect. Veronica subsect. Petraeae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Multiflorae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977, pro parte [only V. umbrosa].

V. sect. Veronica subsect. Acrothecae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977.

V. sect. Veronica subsect. Orientales (E. Wulff) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977.

V. sect. Veronica subsect. Pseudo-orientales Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Thymifolia Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977, pro parte [excl. V. thessalica Benth. in DC.]

V. sect. Veronica subsect. Caespitosa Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Glabrifoliae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Microcarpae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Veronica subsect. Euphrasiifoliae (Römpp) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Armeno-Persicae (Riek.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Veronica subsect sect. Turcomanicae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977.

V. sect. Veronica subsect. Minutae (Jelen.) Mart. Ortega & E. Rico in Bot. Journ. Linn. Soc 135: 185. 2001.[=V. sect. Veronicastrum subsect. Minutae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977, pro parte, excl. V. telephiifolia, V. glareosa, V. chionantha, V. kopetdaghensis et V. violifolia].
 
 

Preliminary list of species:

 Veronica acrotheca, allahuekberensis, altaica, antalyensis, aragonensis, arceuthobia, armena, aucheri, austriaca, baranetzki, bogosensis, (bombycina ssp.) bolkardaghensis, bombycina ssp. bombycina, ssp. froediniana, caespitosa, carduchorum, caucasica, cetikiana, cinerea, crinita, cuneifolia ssp. cuneifolia, ssp. isaurica, ssp. massicytica, czerniakowskiana, denudata, dichrus, elmaliensis, euphrasiifolia, farinosa, filifolia, fontqueri, fragilis, fridericae, fuhsii, galathica, glabrifolia, jacquinii, javalambrensis, khorassanica, kindlii, kopetdaghensis, krylovii, kurdica ssp. filicaulis, ssp. kurdica, leiocarpa, liwanensis, macrostachya ssp. iconiensis, ssp. macrostachya, ssp. mardinensis, ssp. pisidica, ssp. schizostegia, ssp. sorgerae, microcarpa, mirabilis, montbretii, multifida, nigricans, oltensis, orbiculata, orientalis ssp. carduchorum, ssp. nimrodi, ssp. orientalis, orsiniana, paederotae, pectinata, peduncularis, petraea, polifolia, polium, propinqua, prostrata, rechingeri, rhodopaea, rosea, satureiifolia, sennenii, stenobotrys, surculosa, tauricola, tenuifolia, teucrium, thymifolia, thymoides ssp. hasandaghensis, ssp. pseudocinerea, ssp. thymoides, tripartita, turrilliana, umbrosa
 

Veronica subgen. Fruticulosae (Benth.) Martínez-Ortega, Albach & Fischer (ined.) – Type: Veronica fruticulosa L.

= V. [unranked] Fruticulosae Benth. in DC., Prodr.: 480. 1846. [basionym]

= V. subsect. Fruticulosae (Benth.) Stroh in Beih. Bot. Centralbl. 61 B: 388. 1942.

= V. sect. Stenocarpon Boriss.

= V. sect. Veronicastrum W. D. J. Koch, pro parte

= V. sect. Veronica pro parte
 
 

 Comprises: V. sect. Veronicastrum subsect. Gouani (Moretti) Stroh in Beih. Bot. Centralbl. 61 B: 385. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977, pro parte [excl. V. ponae, V. monticola, V. stelleri].

V. sect. Veronicastrum subsect. Macrostemon (Boriss.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Veronicastrum subsect. Densiflorae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 156. 1977.

V. sect. Stenocarpon subsect. Stenocarpon Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977.

V. sect. Stenocarpon subsect. Longibracteatae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.

V. sect. Stenocarpon subsect. Pamiroaltaicae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977 [orthographical error in Jelenevskij (1977): 158 Pamiroalaicae].

V. sect. Veronicastrum subsect. Fruticulosae (Benth.) Stroh in Beih. Bot. Centralbl. 61 B: 388. 1942., includes also V. mampodrensis P. Monts.

V. sect. Veronicastrum subsect. Diffusae (Benth.) Stroh in Beih. Bot. Centralbl. 61 B: 389. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977.

V. sect. Veronicastrum subsect. Saturejoides Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977.

V. sect. Veronica subsect. Thymifolia Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 155. 1977, pro parte [excl. V. polifolia, V. thymifolia et V. rhodopaea].
 
 

 Preliminary list of species:

Veronica cachemirica capitata, chinoalpina, ciliata ssp.cephaloides, ssp.ciliata, ssp.zhongdianensis, densiflora, erinoides, eriogyne, fedtschenkoi, fruticans, fruticulosa, gorbunovii, hirta, kotschyana, lanosa, lanuginosa, lutkeana, macrostemon, macrostemonoides, mampodrensis, mexicana, nummularia, rockii, saturejoides ssp.kellererii, ssp.munellensis, ssp.saturejoides, serpylloides, thessalica
 

Veronica subgen. Chamaedrys (W. D. J. Koch) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. chamaedrys L.

= V. sect. Chamaedrys W. D. J. Koch, Syn. Fl. Germ. Helv.: 524. 1837, pro parte [basionym]

= V. sect. Veronica pro parte

= V. sect. Pocilla Dumort. pro parte

Comprises:

V. sect. Veronica subsect. Multiflorae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977, pro parte [excl. V. umbrosa].

V. sect. Alsinebe subsect. Alsinebe (Griseb.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.
 
 

 Preliminary list of species:

Veronica arvensis, chamaedrys ssp. chamaedrys, ssp. chamaedryoides, ssp. micans, dillenii, laxa, magna, micrantha, sartoriana, verna, vindobonensis
 

Veronica subgen. Pellidosperma (E. B. J. Lehm.) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. praecox All.

= V. [unranked] Pellidosperma E. B. J. Lehm. in Z. Bot. 2: 599. 1910. [basionym]

= V. sect. Alsinebe subsect. Pellidosperma (E. B. J. Lehm.) Stroh in Beih. Bot. Centralbl. 61 B: 399. 1942.

                     = V. sect. Pocilla Dumort., pro parte Comprises:

V. sect. Alsinebe subsect. Pellidosperma (E. B. J. Lehm.). Stroh in Beih. Bot. Centralbl. 61 B: 399. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977, pro parte [excl.. V. bungei]

V. sect. Alsinebe subsect. Amoenae Jelen.in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977, pro parte [excl. V. amoena].
 
 

 Preliminary list of species:

Veronica aznavourii, donii, glauca ssp. chaubardii, ssp. glauca, ssp. peleponnesica, praecox, samuelssonii, triphyllos
 
 

Veronica subgen. Pocilla (Dumort.) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. agrestis L.

= V. [unranked] Pocilla Dumort., Fl. Belg.: 35. 1827 [basionym]

= V. sect. Pocilla pro parte

Comprises:

V. sect. Alsinebe subsect. Brevistylae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977 [excl. V. viscosa].

V. sect. Alsinebe subsect. Biloba (E. B. J. Lehm.) Stroh in Beih. Bot. Centralbl. 61 B: 402. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977.

V. sect. Alsinebe subsect. Tetraphyllae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977.

V. sect. Alsinebe subsect. Cardiocarpae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.

V. sect. Alsinebe subsect. Pellidosperma (E. B. J. Lehm.). Stroh in Beih. Bot. Centralbl. 61 B: 399. 1942, emend. Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977, pro parte [only V. bungei].

V. sect. Alsinebe subsect. Ceratocarpae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977.

V. sect. Alsinebe subsect. Amoenae Jelen.in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 158. 1977, pro parte [only V. amoena].

V. sect. Alsinoides subsect. Filiformes Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977.

V. sect. Alsinoides subsect. Alsinoides (W. D. J. Koch) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 160. 1977.
 
 

 Preliminary list of species:

Veronica agrestis, albanica, amoena, argute-serrata, avromanica, biloba, bucharica, bungei, campylopoda, capillipes, cardiocarpa, ceratocarpa, chantavica, ferganica, filiformis, francispetae, intercedens, karatavica, macropoda, nevskii, opaca, persica, polita, ramosissima, rubrifolia ssp.respectatissima, ssp.rubrifolia, siaretensis, stylophora, tenuissima

Veronica subgen. Digitatae (Jelen.) Martínez-Ortega, Albach & Fischer (ined.) – Type: V. chamaepithyoides Lam.

= V. sect. Alsinebe subsect. Digitatae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977 [basionym]

= V. sect. Pocilla Dumort., pro parte

Comprises:

V. sect. Alsinebe subsect. Digitatae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977.
 
 

 Preliminary list of species:

Veronica chamaepithyoides, grisebachii

Veronica subgen. Veronicella (Heist. ex Fabr.) Pennell in Rhodora 23: 7. 1921, emend. Martínez-Ortega, Albach & Fischer (ined.)., non V. subgen. Veronicella (Fourr.) Boriss. in Shiskhin & Bobrov, Fl. USRR 22: PP. 1955, comb. illeg. – Type: V. hederifolia L.

= Veronicella Heist. ex Fabr., Enum., ed. 1: 111, 1759 [basionym], non Fourr.

= V. [unranked] Cochlidiosperma Rchb., Fl. Germ. Excurs.: 365. 1831-32, pro parte; [=Cochlidiosperma Rchb., Consp. Regn. Veg.: 121. 1828, nom. inval., nom nud.] ; [=Cochlidiosperma (Rchb.) Rchb. Handb. Nat. Pfl.-Syst. 198, 1837], [=V. sect. Cochlidiosperma (Rchb.) Benth., Scroph. Ind.: 45. 1835.]

= V. sect. Pocilla Dumort., pro parte
 
 

 Comprises: V. sect. Diplophyllum subsect. Diplophyllum (E. B. J. Lehm., Ges. Nat. Fr. Berl.Mag. 8: 340-342) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 160. 1977., nom. rej. (art. 56.1, based on Diplophyllum nom. rej.) [= Oligospermum Hong, Opera Bot. 75: 56. 1984] [= Diplophyllum E. B. J. Lehmm., Ges. Nat. Fr. Berl. Mag. 8: 340-342, nom. rej.]

V. sect. Diplophyllum subsect. Cymbalariae (Benth.) Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 160. 1977.
 
 

 Preliminary list of species:

Veronica crista-galli, cymbalaria, hederifolia, lycica, panormitana ssp. baradostensis, ssp. panormitana, sibthorpioides, stamatiadae, stewartii, sublobata, trichadena, triloba
 
 

Veronica subgen. Synthyris (Benth.) Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 87 1933, emend. Martínez-Ortega, Albach & Fischer (ined.). – Type: V. missourica Raf.

=Synthyris Benth. in DC., Prodr. 10: 454. 1846. [basionym]  
=Veronica subgen. Plagiocarpus Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 86. 1933.

= Veronica subgen. Eusynthyris Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 87. 1933, nom. inval.

Comprises:

Besseya Rydb., Bull. Torrey Bot. Club 30: 279. 1903.

Synthyris Benth. in DC., Prodr. 10: 454. 1846
 
 

 Includes (based on species list for Synthyris and Besseya provided by L. Hufford):

V. besseyaMart. Ortega, Albach & M. A. Fisch., nom. nov. pro Besseya alpina (A. Gray) Rydb.

=Synthyris alpina A. Gray in Amer. J. Sci. II. 34: 251. 1862.

=Wulfenia alpina (A. Gray) Greene in Erythea 2: 83. 1894.

=Besseya alpina (A. Gray) Rydb. in Bull. Torrey Bot. Club 30: 279. 1903.

V. bullii (Eaton)Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Gymnandra bullii Eaton in Eaton & Wright, Man. Bot., ed. 8 [North American Botany]: 259. 1840. [basionym]

=Wulfenia bullii (Eaton) Barnhart in Bull. Torrey Bot. Club 26: 378. 1899.

=Synthyris bullii (Eaton) A. Heller in Muhlenbergia 1: 4. 1900.

=Besseya bullii (Eaton) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

= Synthyris houghtoniana Benth., in DC. Prod. 10: 454. 1846.

V. oblongifolia (Pennell) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Besseya oblongifolia Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 101. 1933. [basionym]

V. plantaginea E. James in Trans. Amer. Philos. Soc. II. 2: 173. 1825.

=Synthyris plantaginea (E. James) Benth. in DC., Prod. 10: 455. 1846.

=Wulfenia plantaginea (E. James) Greene in Erythea 2: 83. 1894.

=Besseya plantaginea (E. James) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

= Besseya arizonica Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 103. 1933.

= Besseya goodingii Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 101. 1933

V. ritteriana (Eastw.) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Synthyris ritteriana Eastw. in Proc. Calif. Acad. Sci. II, 1: 123, pl. 9. 1898. [basionym]

=Besseya ritteriana (Eastw.) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

= Synthyris reflexa Eastwood in Proc. Calif. Acad. Sci. II, 1:124, pl. 9. 1898.

= Besseya reflexa (Eastwood) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

= Synthyris flavescens A. Nelson in Bot. Gaz. 34: 33. 1902.

V. rubra (Douglas)Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Gymnandra rubra Douglas in Hook., Fl. Bor.-Amer. 2: 103. 1838. [basionym]

=Synthyris rubra (Douglas) Benth. in DC., Prod. 10: 455. 1846.

=Wulfenia rubra (Douglas) Greene in Erythea 2: 83. 1894.

=Besseya rubra (Douglas) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

=Lunellia rubra (Douglas) Nieuwl. in Amer. Midl. Naturalist 3: 189. 1914.

V. wyomingensis (A. Nelson) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Wulfenia wyomingensis A. Nelson in Bull. Torrey Bot. Club 25: 281. 1898. [basionym]

=Synthyris wyomingensis (A. Nelson) A. Heller in Muhlenbergia 1: 5. 1900.

=Besseya wyomingensis (A. Nelson) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

= Veronica cinerea Raf., New. Fl. 4: 39. 1838, nom. rej. prop., [non V. cinerea Boiss. & Balansa, Diagn. Pl. Orient., Ser. 2, vol. 6: 131. 1859, nom. cons. prop.] (Martínez-Ortega & Albach, same issue).

= Besseya cinerea Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 104. 1933, nom. illeg.

= Wulfenia gymnocarpa A. Nelson in Bull. Torrey Bot. Club 25: 282. 1898.

= Synthyris gymnocarpa (A. Nelson) A. Heller in Muhlenbergia 1: 5. 1900.

= Besseya gymnocarpa (A. Nelson) Rydb. in Bull. Torrey Bot. Club 30: 280. 1903.

V. alaskensis Mart. Ortega, Albach & M. A. Fisch., nom. nov. pro Synthyris borealis Pennell

=Synthyris borealis Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 88. 1933.

V. canbyi (Pennell)Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Synthyris canbyi Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 93. 1933. [basionym]

V. californica Mart. Ortega, Albach & M. A. Fisch., nom. nov. pro Synthyris cordata (A. Gray) Heller

=Synthyris reniformis [unranked] cordata A. Gray, Bot. Calif. 1: 571. 1876.

=Synthyris cordata (A. Gray) A. Heller in Muhlenbergia 1: 4. 1900.

V. dissecta (Rydb.)Mart. Ortega, Albach & M. A. Fisch., comb. nov. var. dissecta

=Synthyris dissecta Rydb. in Bull. Torrey Bot. Club 36: 691. 1909. [basionym]

= Synthyris hendersonii Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 94. 1933.

= Synthyris cymopteroides Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 94. 1933.

V. dissecta var. lanuginosa (Piper) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

= Synthyris pinnatifida [unranked] lanuginosa Piper in Contr. U. S. Natl. Herb. 11: 504. 1906. [basionym]

= Synthyris lanuginosa (Piper) Pennell & J. W. Thomps. in Proc. Acad. Nat. Sci. Philadelphia 85: 93. 1933.

V. utahensis Mart. Ortega, Albach & M. A. Fisch., nom. nov. pro Synthyris laciniata (A. Gray) Rydb.

=Synthyris pinnatifida [unranked] laciniata A. Gray , Syn. Fl. N. Amer. 2, part 1: 286. 1878.

=Synthyris laciniata (A. Gray) Rydb. in Mem. New York Bot. Gard. 1: 353. 1900.

V. missuricaRaf. in Am. Monthly Mag. & Crit. Rev. 3: 175. 1818. ssp. missurica

=V. reniformis Pursh, Fl. Amer. Sept. 1: 10. 1814, nom. illeg. [non Veronica reniformis Raf. in Med. Repos. 5: 360. 1808.]

=V. purshii G. Don, Gen. Hist. 4: 573. 1838.

=Synthyris missurica (Raf.) Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 89. 1933.

= Synthyris missurica [unranked] hirsuta Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 89. 1933.

V. missuricaRaf. ssp. major(Hook.) Mart. Ortega, Albach & M. A. Fisch.,comb. nov.

=Synthyris reniformis ssp. major Hook. in J. Bot. (Hooker) 5: 257. 1853. [basionym]

=Synthyris major (Hook.) A. Heller in Muhlenbergia 1: 4. 1900.

V. missuricaRaf. ssp. stellata(Pennell) Mart. Ortega, Albach & M. A. Fisch.,comb. nov.

=Synthyris stellata Pennell in Proc. Acad. Nat. Sci. Philadelphia 85. 89. 1933. [basionym]

V. paysoni (Pennell & L. Williams) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Synthyris paysoni Pennell & L. Williams in Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 96. 1933. [basionym]

= Synthyris pinnatifida S. Watson, Botany [fortieth parallel]: 227, pl. 22. 1871.

V. idahoensis (Gail & Pennell) Mart. Ortega, Albach & M. A. Fisch., nom. nov. pro Synthyris platycarpa Gail & Pennell

=Synthyris platycarpa Gail & Pennell in Amer. J. Bot. 24: 40. 1937.

V. ranunculina (Pennell) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Synthyris ranunculina Pennell in Proc. Acad. Nat. Sci. Philadelphia 85: 92. 1933. [basionym]

V. regina-nivalis Mart. Ortega, Albach & M. A. Fisch., nom. nov. pro Synthyris reniformis (Douglas) Benth. [Typus subgeneris]

=Wulfenia reniformis Douglas ex Benth., Scroph. Ind.: 46. 1835.

=Synthyris reniformis (Douglas ex Benth.) Benth. in DC., Prodr. 10: 454. 1846.

= Synthyris rotundifolia A. Gray, Syn. Fl. N. Amer. 2, part 1: 285. 1878.

V. schizantha (Piper) Mart. Ortega, Albach & M. A. Fisch., comb. nov.

=Synthyris schizantha Piper in Bull. Torr. Bot. Club 29: 223. 1902. [basionym]
 

VERONICA INCERTA SEDE

V. sect. Beccabunga subsect. Himalensis Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 152. 1977.

V. sect. Veronica subsect. JavanicaeJelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 154. 1977, pro parte [excl. V. morrisonicola].

V. sect. Veronicastrum subsect. Alpinae (Benth.) Stroh in Beih. Bot. Centralbl. 61 B: 386. 1942., pro parte [only V. chinoalpina]

V. sect. Veronicastrum subsect. Tianschanicae Boriss. ex Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977.

V. sect. Veronica subsect. Minutae (Jelen.) Mart. Ortega & E. Rico in Bot. Journ. Linn. Soc 135: 185. 2001.[=V. sect. Veronicastrum subsect. Minutae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 157. 1977, pro parte [only V. chionantha, V. kopetdaghensis et V. violifolia].

V. sect. Alsinebe subsect. Brevistylae Jelen. in Byull. Moskovsk. Obshch. Isp. Prir. Otd. Biol. 82 (1): 159. 1977 [only V. viscosa].

V. sect. Paederotella E. Wulff in Trudy Tiflissk. Bot. Sada 15: 68. 1915. - Type: V. ruprechtii Lipsky. [V. subgen. Paederotella (E. Wulff) Boriss. in Shiskhin & Bobrov, Fl. USRR 22: 492. 1955.]

                     V. sect. Paederotoides Benth. in DC., Prodr.: 479. 1846.